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Introduction

The MuData format (.h5mu) stores multimodal single-cell data in a single file. Each modality (RNA, protein, ATAC) lives as a separate AnnData object, sharing a common set of cell barcodes. This is the native format for muon in Python. scConvert reads and writes h5mu natively – no Python or MuDataSeurat required.

Format Best for
h5ad Single-modality (RNA only)
h5mu Multi-modal (RNA + ADT, RNA + ATAC, etc.)

Read h5mu directly

The shipped citeseq_demo.h5mu contains 500 CITE-seq cells with two modalities: RNA (2,000 genes) and ADT (10 surface protein antibodies). readH5MU() loads it into a Seurat object with both assays intact.

h5mu_file <- system.file("extdata", "citeseq_demo.h5mu", package = "scConvert")
obj <- readH5MU(h5mu_file)

cat("Cells:", ncol(obj), "\n")
#> Cells: 500
cat("Assays:", paste(Assays(obj), collapse = ", "), "\n")
#> Assays: ADT, RNA
cat("RNA features:", nrow(obj[["RNA"]]), "\n")
#> RNA features: 2000
cat("ADT features:", nrow(obj[["ADT"]]), "\n")
#> ADT features: 10

UMAP of RNA clusters

DimPlot(obj, group.by = "seurat_clusters", label = TRUE, pt.size = 0.8) +
  ggtitle("CITE-seq clusters (loaded from h5mu)")

Protein expression

After reading h5mu, the ADT assay contains only raw counts. Normalize with CLR (centered log-ratio) before plotting protein markers.

DefaultAssay(obj) <- "ADT"
obj <- NormalizeData(obj, normalization.method = "CLR", margin = 2, verbose = FALSE)
FeaturePlot(obj, features = "CD3", pt.size = 0.8) +
  ggtitle("CD3 protein (ADT) -- loaded from h5mu")

DefaultAssay(obj) <- "RNA"

Write and roundtrip

Load the same dataset from its Seurat .rds form, write to h5mu, then read back and compare. This demonstrates that scConvert preserves both assays through a full write/read cycle.

Modality name mapping

writeH5MU() automatically maps Seurat assay names to standard MuData conventions:

Seurat Assay h5mu Modality
RNA rna
ADT prot
ATAC atac
Other lowercase name

readH5MU() reverses the mapping when loading.

orig <- readRDS(system.file("extdata", "citeseq_demo.rds", package = "scConvert"))

h5mu_path <- file.path(tempdir(), "citeseq_roundtrip.h5mu")
writeH5MU(orig, h5mu_path, overwrite = TRUE)
cat("Wrote:", round(file.size(h5mu_path) / 1024^2, 1), "MB\n")
#> Wrote: 0.7 MB

loaded <- readH5MU(h5mu_path)
cat("Loaded:", ncol(loaded), "cells,", paste(Assays(loaded), collapse = ", "), "\n")
#> Loaded: 500 cells, ADT, RNA

Side-by-side comparison

library(patchwork)

p1 <- DimPlot(orig, group.by = "seurat_clusters", label = TRUE, pt.size = 0.8) +
  ggtitle("Original (.rds)") + NoLegend()
p2 <- DimPlot(loaded, group.by = "seurat_clusters", label = TRUE, pt.size = 0.8) +
  ggtitle("After h5mu roundtrip") + NoLegend()
p1 + p2

Verify data integrity

cat("Cell count match:", ncol(orig) == ncol(loaded), "\n")
#> Cell count match: TRUE
cat("Assays match:", identical(sort(Assays(orig)), sort(Assays(loaded))), "\n")
#> Assays match: TRUE

common_cells <- intersect(colnames(orig), colnames(loaded))
common_genes <- intersect(rownames(orig[["RNA"]]), rownames(loaded[["RNA"]]))
orig_rna <- as.numeric(GetAssayData(orig, assay = "RNA", layer = "counts")[
  head(common_genes, 100), head(common_cells, 100)])
rt_rna <- as.numeric(GetAssayData(loaded, assay = "RNA", layer = "counts")[
  head(common_genes, 100), head(common_cells, 100)])
cat("RNA counts identical:", identical(orig_rna, rt_rna), "\n")
#> RNA counts identical: TRUE

common_adt <- intersect(rownames(orig[["ADT"]]), rownames(loaded[["ADT"]]))
orig_adt <- as.numeric(GetAssayData(orig, assay = "ADT", layer = "counts")[
  common_adt, head(common_cells, 100)])
rt_adt <- as.numeric(GetAssayData(loaded, assay = "ADT", layer = "counts")[
  common_adt, head(common_cells, 100)])
cat("ADT counts identical:", identical(orig_adt, rt_adt), "\n")
#> ADT counts identical: TRUE

Custom modality name mapping

You can override the default mapping when reading with the assay.names argument:

custom <- readH5MU(h5mu_file, assay.names = c(rna = "RNA", prot = "Protein"))
cat("Assays with custom mapping:", paste(Assays(custom), collapse = ", "), "\n")
#> Assays with custom mapping: Protein, RNA

Python interoperability

The h5mu file is directly compatible with muon in Python.

# Requires Python: pip install mudata
import mudata as md

mdata = md.read_h5mu("citeseq_demo.h5mu")
print(mdata)
print("Modalities:", list(mdata.mod.keys()))
for name, mod in mdata.mod.items():
    print(f"  {name}: {mod.n_obs} cells x {mod.n_vars} features")

Clean up

unlink(h5mu_path)

Session Info

sessionInfo()
#> R version 4.5.2 (2025-10-31)
#> Platform: aarch64-apple-darwin20
#> Running under: macOS Tahoe 26.3
#> 
#> Matrix products: default
#> BLAS:   /System/Library/Frameworks/Accelerate.framework/Versions/A/Frameworks/vecLib.framework/Versions/A/libBLAS.dylib 
#> LAPACK: /Library/Frameworks/R.framework/Versions/4.5-arm64/Resources/lib/libRlapack.dylib;  LAPACK version 3.12.1
#> 
#> locale:
#> [1] en_US.UTF-8/en_US.UTF-8/en_US.UTF-8/C/en_US.UTF-8/en_US.UTF-8
#> 
#> time zone: America/Indiana/Indianapolis
#> tzcode source: internal
#> 
#> attached base packages:
#> [1] stats     graphics  grDevices utils     datasets  methods   base     
#> 
#> other attached packages:
#> [1] patchwork_1.3.2    ggplot2_4.0.2      Seurat_5.4.0       SeuratObject_5.3.0
#> [5] sp_2.2-1           scConvert_0.1.0   
#> 
#> loaded via a namespace (and not attached):
#>   [1] RColorBrewer_1.1-3     jsonlite_2.0.0         magrittr_2.0.4        
#>   [4] spatstat.utils_3.2-2   farver_2.1.2           rmarkdown_2.30        
#>   [7] fs_1.6.7               ragg_1.5.0             vctrs_0.7.1           
#>  [10] ROCR_1.0-12            spatstat.explore_3.7-0 htmltools_0.5.9       
#>  [13] sass_0.4.10            sctransform_0.4.3      parallelly_1.46.1     
#>  [16] KernSmooth_2.23-26     bslib_0.10.0           htmlwidgets_1.6.4     
#>  [19] desc_1.4.3             ica_1.0-3              plyr_1.8.9            
#>  [22] plotly_4.12.0          zoo_1.8-15             cachem_1.1.0          
#>  [25] igraph_2.2.2           mime_0.13              lifecycle_1.0.5       
#>  [28] pkgconfig_2.0.3        Matrix_1.7-4           R6_2.6.1              
#>  [31] fastmap_1.2.0          fitdistrplus_1.2-6     future_1.69.0         
#>  [34] shiny_1.13.0           digest_0.6.39          tensor_1.5.1          
#>  [37] RSpectra_0.16-2        irlba_2.3.7            textshaping_1.0.4     
#>  [40] labeling_0.4.3         progressr_0.18.0       spatstat.sparse_3.1-0 
#>  [43] httr_1.4.8             polyclip_1.10-7        abind_1.4-8           
#>  [46] compiler_4.5.2         bit64_4.6.0-1          withr_3.0.2           
#>  [49] S7_0.2.1               fastDummies_1.7.5      MASS_7.3-65           
#>  [52] tools_4.5.2            lmtest_0.9-40          otel_0.2.0            
#>  [55] httpuv_1.6.16          future.apply_1.20.2    goftest_1.2-3         
#>  [58] glue_1.8.0             nlme_3.1-168           promises_1.5.0        
#>  [61] grid_4.5.2             Rtsne_0.17             cluster_2.1.8.2       
#>  [64] reshape2_1.4.5         generics_0.1.4         hdf5r_1.3.12          
#>  [67] gtable_0.3.6           spatstat.data_3.1-9    tidyr_1.3.2           
#>  [70] data.table_1.18.2.1    spatstat.geom_3.7-0    RcppAnnoy_0.0.23      
#>  [73] ggrepel_0.9.7          RANN_2.6.2             pillar_1.11.1         
#>  [76] stringr_1.6.0          spam_2.11-3            RcppHNSW_0.6.0        
#>  [79] later_1.4.8            splines_4.5.2          dplyr_1.2.0           
#>  [82] lattice_0.22-9         survival_3.8-6         bit_4.6.0             
#>  [85] deldir_2.0-4           tidyselect_1.2.1       miniUI_0.1.2          
#>  [88] pbapply_1.7-4          knitr_1.51             gridExtra_2.3         
#>  [91] scattermore_1.2        xfun_0.56              matrixStats_1.5.0     
#>  [94] stringi_1.8.7          lazyeval_0.2.2         yaml_2.3.12           
#>  [97] evaluate_1.0.5         codetools_0.2-20       tibble_3.3.1          
#> [100] cli_3.6.5              uwot_0.2.4             xtable_1.8-8          
#> [103] reticulate_1.45.0      systemfonts_1.3.1      jquerylib_0.1.4       
#> [106] dichromat_2.0-0.1      Rcpp_1.1.1             globals_0.19.1        
#> [109] spatstat.random_3.4-4  png_0.1-8              spatstat.univar_3.1-6 
#> [112] parallel_4.5.2         pkgdown_2.2.0          dotCall64_1.2         
#> [115] listenv_0.10.1         viridisLite_0.4.3      scales_1.4.0          
#> [118] ggridges_0.5.7         purrr_1.2.1            crayon_1.5.3          
#> [121] rlang_1.1.7            cowplot_1.2.0